Anaerobic microorganisms play a significant role in global cycling of elements and nutrients and play critical roles in health and disease processes in animals and humans [[1], [2], [3], [4], [5]]. In addition, many anaerobes are used in industrial and commercial applications [[6], [7], [8], [9]]. Strictly anaerobic microorganisms are predominant members of a wide range of anoxic ecosystems such as freshwater and marine sediments and soils, and the gastrointestinal tracts of insects, birds,