Kharadi, A., Patel, M., & Patel, F. 2019. Assessment of extra cellular enzymes of bacteria isolated from mangrove rhizosphere soil of different places of Gujarat in monsoon season. Indian Journal of Science and Technology, 12, 45.‏

Muñoz-García, A., Arbeli, Z., Boyacá-Vásquez, V., & Vanegas, J. 2022. Metagenomic and genomic characterization of heavy metal tolerance and resistance genes in the rhizosphere microbiome of Avicennia germinans in a semi-arid mangrove forest in the tropics. Marine Pollution Bulletin, 184, 114204.

Samara, F., Solovieva, N., Ghalayini, T., Nasrallah, Z., & Saburova, M. 2020. Assessment of the environmental status of the mangrove ecosystem in the United Arab Emirates. Water, 12, 1623.

Alzahrani, D., Selim, E.-M., & El-Sherbiny, M. 2018. Ecological assessment of heavy metals in the grey mangrove (Avicennia marina) and associated sediments along the Red Sea coast of Saudi Arabia. Oceanologia, 60, 513-526.

Roy, R., Samanta, S., Pandit, S., Naaz, T., Banerjee, S., Rawat, J., ... & Saha, R. 2024. An overview of bacteria-mediated heavy metal bioremediation strategies. Applied Biochemistry and Biotechnology, 196, 1712-1751.

John, C., ChukwumatiJohn, A., & George, A. 2023. Accumulation of hydrocarbons and some heavy metal contents on sediments and plants from crude oil polluted mangrove ecosystem in Okrika, Nigeria. World Journal of Advanced Research and Reviews, 17, 298-306.

Goyal, P., Belapurkar, P., & Kar, A. 2019. A review on in vitro and in vivo bioremediation potential of environmental and probiotic species of Bacillus and other probiotic microorganisms for two heavy metals, cadmium and nickel. Biosciences Biotechnology Research Asia, 16, 01-13.

Saini, S., & Dhania, G. 2020. Cadmium as an environmental pollutant: ecotoxicological effects, health hazards, and bioremediation approaches for its detoxification from contaminated sites. Bioremediation of industrial waste for environmental safety: Volume II: biological agents and methods for industrial waste management. Singapore: Springer Singapore, 357-387.

Sekar, A., & Kim, K. 2020. Halophilic bacteria in the food industry. Encyclopedia of Marine Biotechnology, 4, 2061-2070.

Biswas, J., Jana, S., & Mandal, S. 2023. Biotechnological impacts of Halomonas: a promising cell factory for industrially relevant biomolecules. Biotechnology and Genetic Engineering Reviews, 39, 348-377.

Lata, S., Kaur, H., & Mishra, T. 2019. Cadmium bioremediation: a review. International Journal of Pharmaceutical Sciences and Research, 10, 4120-4128.

Zmorrod, Z., Nadine, G., Al Hakawati, N., Amer, R., & Yusef, H. 2021. Lead and cadmium detoxification by halophilic bacteria isolated from solar salterns in Lebanon. BAU Journal-Science and Technology, 2, 3.

Baati, H., Bahloul, M., Amdouni, R., & Azri, C. 2022. Behavior assessment of moderately halophilic bacteria in brines highly enriched with heavy metals: Sfax solar saltern (Tunisia), a Case Study. Geomicrobiology Journal, 39, 341-351.

Gan, L., Zhang, H., Tian, J., Li, X., Long, X., Zhang, Y., ... & Tian, Y. 2018. Planococcus salinus sp. nov., a moderately halophilic bacterium isolated from a saline-alkali soil. International Journal of Systematic and Evolutionary Microbiology, 68, 589-595.

Elyasifar, B., Jafari, S., Hallaj-Nezhadi, S., Chapeland-leclerc, F., Ruprich-Robert, G., & Dilmaghani, A. 2019. Isolation and identification of antibiotic-producing halophilic bacteria from Dagh Biarjmand and Haj Aligholi salt deserts, Iran. Pharmaceutical Sciences, 25, 70-77.

Nieto, J., Fernandez-Castillo, R., Marquez, M., Ventosa, A., Quesada, E., & Ruiz-Berraquero, F. 1989. Survey of metal tolerance in moderately halophilic eubacteria. Applied and Environmental Microbiology, 55, 2385-2390.

Al-Dhabi, N., Esmail, G., Mohammed Ghilan, A.-K., & Valan Arasu, M. 2019. Optimizing the management of cadmium bioremediation capacity of metal-resistant Pseudomonas sp. strain Al-Dhabi-126 isolated from the industrial city of Saudi Arabian environment. International Journal of Environmental Research and Public Health, 16, 4788.

Orji, O., Awoke, J., Aja, P., Aloke, C., Obasi, O., Alum, E., ... & Oka, G. 2021. Halotolerant and metalotolerant bacteria strains with heavy metals biorestoration possibilities isolated from Uburu Salt Lake, Southeastern, Nigeria. Heliyon, 7.

Sachindra, N., Sato, E., Maeda, H., Hosokawa, M., Niwano, Y., Kohno, M., & Miyashita, K. 2007. Radical scavenging and singlet oxygen quenching activity of marine carotenoid fucoxanthin and its metabolites. Journal of Agricultural and Food Chemistry, 55, 8516-8522.

Mohapatra, R., Parhi, P., Pandey, S., Bindhani, B., Thatoi, H., & Panda, C. 2019. Active and passive biosorption of Pb (II) using live and dead biomass of marine bacterium Bacillus xiamenensis PbRPSD202: Kinetics and isotherm studies. Journal of Environmental Management, 247, 121-134.‏

Sowmya, M., Rejula, M., Rejith, P., Mohan, M., Karuppiah, M., & Hatha, A. 2014. Heavy metal tolerant halophilic bacteria from Vembanad Lake as possible source for bioremediation of lead and cadmium. Journal of Environmental Biology, 35, 655.

Divakar, G., Sameer, R., & Bapuji, M. 2018. Screening of multi-metal tolerant halophilic bacteria for heavy metal remediation. International Journal of Current Microbiology and Applied Sciences, 7, 2062-2076.

Mgbodile, C., Otutu, U., Onuoha, S., Eze, U., Ugwuoji, T., Nnabuife, O., & Nwagu, T. 2022. Hydrolases secreting, heavy metal-resistant halophilic bacteria isolated from metal dumpsites. Asian Journal of Tropical Biotechnology, 19.

Statistics, L. 2015. Kaplan-Meier using SPSS statistics. Statistical tutorials and software guides.

Chang, H., Han, S., Kim, S., An, J., Alatalo, J., & Son, Y. 2020. Interactions between topsoil properties and ecophysiological responses of mangroves (Avicennia marina) along the tidal gradient in an arid region in Qatar. Turkish Journal of Agriculture and Forestry, 44, 121-126.

Amasha, R., & Alhazzani, R. 2025. Lead bioremoval by halotolerant bacteria isolated from mangrove rhizosphere soils of Saudi Arabia. Journal of King Abdulaziz University: Marine Science, 35, 138-147.‏

Kannika, C., & Thalisa, Y.-A. 2021. The diversity of halotolerant and halophilic bacteria in the soil of the Nasinuan secondary forest in Maha Sarakham, Thailand. Journal of Sustainability Science and Management, 16, 165-175.‏

Chaida, A., Bensalah, F., & M'rassi, A. 2021. Isolation and characterisation of a new alkali-halotolerant Bacillus aquimaris strain lgmt10, producing extracellular hydrolases, from the effluents of a thermal power plant, in Algeria. Journal of Microbiology, Biotechnology and Food Sciences, 11, e3460-e3460.

Claros, A., Guardia, E., & Ayala-Borda, P. 2023. Characterization and heavy metal bioremediation potential of Halomonas isolates from the Bolivian Altiplano.‏ International Journal of Ecotoxicology and Ecobiology, 8, 13-23.

Amoozegar, M., Hamedi, J., Dadashipour, M., & Shariatpanahi, S. 2005. Effect of salinity on the tolerance to toxic metals and oxyanions in native moderately halophilic spore-forming bacilli. World Journal of Microbiology and Biotechnology, 21, 1237-1243.

Castillo-Zacarías, C., Suárez-Herrera, M., Garza-González, M., Sánchez-González, M., & López-Chuken, U. 2011. Cadmium increases catechol 2, 3-dioxygenase activity in Variovorax sp. 12S, a metaltolerant and phenol-degrading strain. African Journal of Microbiology Research, 5, 2627-2631.

Sahoo, S., & Goli, D. (2020). Bioremediation of lead by a halophilic bacteria Bacillus pumilus isolated from the mangrove regions of Karnataka. International Journal of Science and Research, 9, 1337-1343.

Mohapatra, R., Parhi, P., Patra, J., Panda, C., & Thatoi, H. 2018. Biodetoxification of toxic heavy metals by marine metal resistant bacteria - a novel approach for bioremediation of the polluted saline environment. In Microbial Biotechnology: Volume 1. Applications in Agriculture and Environment, 343-376. Singapore: Springer Singapore.‏

Di Cerbo, A., Palmieri, B., Aponte, M., Morales-Medina, J., & Iannitti, T. 2016. Mechanisms and therapeutic effectiveness of lactobacilli. Journal of Clinical Pathology, 69, 187-203.

El Far, M., Zakaria, A., Kassem, M., Wedn, A., Guimei, M., & Edward, E. 2023. Promising biotherapeutic prospects of different probiotics and their derived postbiotic metabolites: in-vitro and histopathological investigation. BMC Microbiology, 23, 122.‏

Neagu, S., & Stancu, M. 2025. Novel halotolerant bacteria from saline environments: isolation and biomolecule production. BioTech, 14, 49.‏

Perumal K., Seenuvasan, J., Nandhagopal, M. & Manivannan, N. 2024. Antimicrobial properties of secondary metabolites produced by Halomonas sp.: a halophilic bacterium. Cureus, 16.

Bahamdain, L., & Aly, M. 2026. A new source of actinomycetes with heavy metal tolerance and antibacterial properties against multidrug-resistant Staphylococcus hominis. Journal of Contemporary Medical Sciences, 12, 58-67.‏

Gómez-Villegas, P., Vigara, J., Vila, M., Varela, J., Barreira, L., & Léon, R. 2020. Antioxidant, antimicrobial, and bioactive potential of two new haloarchaeal strains isolated from odiel salterns (Southwest Spain). Biology, 9, 298.‏

Squillaci, G., Parrella, R., Carbone, V., Minasi, P., La Cara, F., & Morana, A. 2017. Carotenoids from the extreme halophilic archaeon Haloterrigena turkmenica: Identification and antioxidant activity. Extremophiles, 21, 933-945.

Hou, J., & Cui, H. 2018. In vitro antioxidant, antihemolytic, and anticancer activity of the carotenoids from halophilic archaea. Current Microbiology, 75, 266-271.‏

Jaswir, I., Noviendri, D., Hasrini, R., & Octavianti, F. 2011. Carotenoids: sources, medicinal properties and their application in food and nutraceutical industry. Journal of Medicinal Plants Research, 5, 7119-7131.‏

Chaari, M., Theochari, I., Papadimitriou, V., Xenakis, A., & Ammar, E. 2018. Encapsulation of carotenoids extracted from halophilic Archaea in oil-in-water (O/W) micro-and nano-emulsions. Colloids and Surfaces B: Biointerfaces, 161, 219-227.‏