AmphibiaWeb (2024) Hyla chrysoscelis: Cope's Gray Treefrog <https://amphibiaweb.org/species/768>. University of California. Accessed March 28 2026

Arch VS, Grafe TU, Narins PM (2008) Ultrasonic signalling by a Bornean frog. Biol Lett 4(1):19–22. https://doi.org/10.1098/rsbl.2007.0494

Article  PubMed  PubMed Central  Google Scholar 

Arch VS, Grafe TU, Gridi-Papp M, Narins PM (2009) Pure ultrasonic communication in an endemic Bornean frog. PLoS ONE 4(4):e5413. https://doi.org/10.1371/journal.pone.0005413

Article  CAS  PubMed  PubMed Central  Google Scholar 

Bates D, Mächler M, Bolker BM, Walker SC (2015) Fitting linear mixed-effects models using lme4. J Stat Soft 67(1):1–48. https://doi.org/10.18637/jss.v067.i01

Baugh AT, Freiler MK, Halstead LN, Kozak K, Bee MA (2026) Testosterone and estradiol predict male calling performance, but not performance-related tradeoffs, in competitive signaling environments in Cope’s gray treefrogs (Hyla chrysoscelis). J Comp Physiol A

Bee MA (2007) Sound source segregation in grey treefrogs: spatial release from masking by the sound of a chorus. Anim Behav 74(3):549–558. https://doi.org/10.1016/j.anbehav.2006.12.012

Article  Google Scholar 

Bee MA (2008a) Finding a mate at a cocktail party: spatial release from masking improves acoustic mate recognition in grey treefrogs. Anim Behav 75(5):1781–1791. https://doi.org/10.1016/j.anbehav.2007.10.032

Article  PubMed  PubMed Central  Google Scholar 

Bee MA (2008b) Parallel female preferences for call duration in a diploid ancestor of an allotetraploid treefrog. Anim Behav 76(3):845–853. https://doi.org/10.1016/j.anbehav.2008.01.029

Article  PubMed  PubMed Central  Google Scholar 

Bee MA (2012) Sound source perception in anuran amphibians. Curr Opin Neurobiol 22(2):301–310. https://doi.org/10.1016/j.conb.2011.12.014

Article  CAS  PubMed  PubMed Central  Google Scholar 

Bee MA (2015) Treefrogs as animal models for research on auditory scene analysis and the cocktail party problem. Int J Psychophysiol 95(2):216–237. https://doi.org/10.1016/j.ijpsycho.2014.01.004

Article  PubMed  Google Scholar 

Bee MA, Christensen-Dalsgaard J (2016) Sound source localization and segregation with internally coupled ears: the treefrog model. Biol Cybern 110(4–5):271–290. https://doi.org/10.1007/s00422-016-0695-5

Article  PubMed  PubMed Central  Google Scholar 

Bee MA, Micheyl C (2008) The cocktail party problem: what is it? How can it be solved? And why should animal behaviorists study it? J Comp Psychol 122(3):235–251. https://doi.org/10.1037/0735-7036.122.3.235

Article  PubMed  PubMed Central  Google Scholar 

Bee MA, Schwartz JJ (2009) Behavioral measures of signal recognition thresholds in frogs in the presence and absence of chorus-shaped noise. J Acoust Soc Am 126(5):2788–2801. https://doi.org/10.1121/1.3224707

Article  PubMed  PubMed Central  Google Scholar 

Bee MA, Vélez A (2018) Masking release in temporally fluctuating noise depends on comodulation and overall level in Cope’s gray treefrog. J Acoust Soc Am 144(4):2354–2362. https://doi.org/10.1121/1.5064362

Article  PubMed  PubMed Central  Google Scholar 

Bee MA, Buschermöhle M, Klump GM (2007) Detecting modulated signals in modulated noise: II. Neural thresholds in the songbird forebrain. Eur J Neurosci 26:1979–1994. https://doi.org/10.1111/j.1460-9568.2007.05805.x

Article  PubMed  Google Scholar 

Booker WW, Gerhardt HC, Lemmon AR, Ptacek MB, Hassinger ATB, Schul J, Lemmon EM (2022) The complex history of genome duplication and hybridization in North American gray treefrogs. Mol Biol Evol 39(2):msab316. https://doi.org/10.1093/molbev/msab316

Bradbury JW, Vehrencamp SL (2011) Principles of animal communication, 2nd edn. Sinauer Associates, Sunderland, MA

Google Scholar 

Branstetter BK, Finneran JJ (2008) Comodulation masking release in bottlenose dolphins (Tursiops truncatus). J Acoust Soc Am 124(1):625–633. https://doi.org/10.1121/1.2918545

Article  PubMed  Google Scholar 

Brenowitz EA (1982) The active space of red-winged blackbird song. J Comp Physiol 147(4):511–522. https://doi.org/10.1007/BF00612017

Article  Google Scholar 

Bronkhorst AW (2000) The cocktail party phenomenon: a review of research on speech intelligibility in multiple-talker conditions. Acustica 86(1):117–128. http://pascal-francis.inist.fr/vibad/index.php?action=getRecordDetail&idt=5720732

Brumm H (ed) (2013) Animal communication and noise. Springer, New York

Google Scholar 

Brumm H, Naguib M (2009) Environmental acoustics and the evolution of bird song. Adv Stud Behav 40:1–33. https://doi.org/10.1016/S0065-3454(09)40001-9

Article  Google Scholar 

Brumm H, Slabbekoorn H (2005) Acoustic communication in noise. Adv Stud Behav 35:151–209. https://doi.org/10.1016/S0065-3454(05)35004-2

Article  Google Scholar 

Brumm H, Todt D (2002) Noise-dependent song amplitude regulation in a territorial songbird. Anim Behav 63(5):891–897. https://doi.org/10.1006/anbe.2001.1968

Article  Google Scholar 

Brumm H, Zollinger SA (2011) The evolution of the Lombard effect: 100 years of psychoacoustic research. Behav 148(11–13):1173–1198. https://doi.org/10.1163/000579511x605759

Article  Google Scholar 

Brumm H, Voss K, Kollmer I, Todt D (2004) Acoustic communication in noise: regulation of call characteristics in a New World monkey. J Exp Biol 207(3):443–448. https://doi.org/10.1242/jeb.00768

Article  PubMed  Google Scholar 

Brush JS, Narins PM (1989) Chorus dynamics of a neotropical amphibian assemblage: comparison of computer-simulation and natural behavior. Anim Behav 37:33–44. https://doi.org/10.1016/0003-3472(89)90004-3

Article  Google Scholar 

Buus S (1985) Release from masking caused by envelope fluctuations. J Acoust Soc Am 78(6):1958–1965. https://doi.org/10.1121/1.392652

Article  CAS  PubMed  Google Scholar 

Buss E, Hall JW, III, Grose JH (2003) The masking level difference for signals placed in masker envelope minima and maxima. J Acoust Soc Am 114(3):1557–1564. https://https://doi.org/10.1121/1.1598199

Caldwell MS, Lee N, Bee MA (2016) Inherent directionality determines spatial release from masking at the tympanum in a vertebrate with internally coupled ears. J Assoc Res Otolaryngol 17(4):259–270. https://doi.org/10.1007/s10162-016-0568-6

Article  PubMed  PubMed Central  Google Scholar 

Caldwell MS, Bee MA (2014) Spatial hearing in Cope's gray treefrog: I. Open and closed loop experiments on sound localization in the presence and absence of noise. J Comp Physiol A 200(4):265–284. https://doi.org/10.1007/s00359-014-0882-6

Cherry EC (1953) Some experiments on the recognition of speech, with one and with two ears. J Acoust Soc Am 25(5):975–979. https://doi.org/10.1121/1.1907229

Article  Google Scholar 

Cooke M (2006) A glimpsing model of speech perception in noise. J Acoust Soc Am 119(3):1562–1573. https://doi.org/10.1121/1.2166600

Article  PubMed  Google Scholar 

Dawkins R, Krebs JR (1978) Animal signals: information or manipulation. In: Krebs JR, Davies NB (eds) Behavioural ecology: an evolutionary approach, vol 2. Blackwell Scientific, Oxford, pp 282–309

Google Scholar 

Dent ML, McClaine EM, Best V, Ozmeral E, Narayan R, Gallun FJ, Sen K, Shinn-Cunningham BG (2009) Spatial unmasking of birdsong in zebra finches (Taeniopygia guttata) and budgerigars (Melopsittacus undulatus). J Comp Psychol 123(4):357–367. https://doi.org/10.1037/a0016898

Article  PubMed  PubMed Central  Google Scholar 

Diepenbrock JP, Jeschke M, Ohl FW, Verhey JL (2017) Comodulation masking release in the inferior colliculus by combined signal enhancement and masker reduction. J Neurophysiol 117(2):853–867. https://doi.org/10.1152/jn.00191.2016

Article  PubMed  Google Scholar 

Dunia R, Narins PM (1989) Tone-derived vs. tone-in-noise-derived filter functions of frog auditory nerve fibers: a comparison. Hear Res 37(3):241–254. https://doi.org/10.1016/0378-5955(89)90025-7

Epp B, Verhey JL (2009a) Combination of masking releases for different center frequencies and masker amplitude statistics. J Acoust Soc Am 126(5):2479–2489. https://doi.org/10.1121/1.3205404

Article  PubMed  Google Scholar 

Epp B, Verhey JL (2009b) Superposition of masking releases. J Comp Neurosci 26(3):393–407. https://doi.org/10.1007/s10827-008-0118-2

Article  Google Scholar 

Fay RR (2011) Signal-to-noise ratio for source determination and for a comodulated masker in goldfish, Carassius auratus. J Acoust Soc Am 129(5):3367–3372. https://doi.org/10.1121/1.3562179

Article  PubMed  PubMed Central  Google Scholar